Kissing feels so tied to human life that it can seem almost impossible to separate from romance, family, grief, ritual, or desire. It sits inside wedding vows and movie endings, in greetings, apologies, and quiet moments that do not need words. Yet mouth-to-mouth contact did not begin with people, and it does not belong to us alone. Across the animal kingdom, behaviors that look a lot like kissing appear in species as different as bonobos, prair…
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