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Gramma the Galapagos Tortoise, Oldest Resident of San Diego Zoo, Dies at About 141

Gramma was euthanized due to age-related bone deterioration after living about 141 years and serving as an ambassador for reptile conservation at the San Diego Zoo.

  • On Thursday, the San Diego Zoo Wildlife Alliance announced Gramma, a Galápagos tortoise, died after care teams euthanized her due to worsening age-related bone deterioration.
  • Arriving between 1928 and 1931, Gramma joined the San Diego Zoo as part of its first Galápagos tortoise group after being collected from the Galápagos Islands in 1835.
  • Experts estimate Gramma may have lived up to 141 years, making her the zoo's oldest resident; zoo officials said her life spanned 20 U.S. presidents, two world wars and two pandemics.
  • The zoo posted video showing Gramma meandering in her habitat and invited members of the public and visitors to honor her with a generous, fruit-filled salad tribute.
  • Conservation groups note that Galápagos tortoises can live over 100 years, include 15 subspecies with three extinct, and conservation programs have released more than 10,000 juveniles since 1965.
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The animal suffered from a bone problem and had to be euthanized.

·Paris, France
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The community of the San Diego Zoo and wildlife lovers from all over the world lament the death of Gramma, an emblematic Galapagos turtle that, at 141 years old, had become one of the most beloved and long-lived animals of the enclosure. Known affectionately as the “reign of the zoo”, Gramma was for decades a symbol of resilience, serenity and connection with the natural history of the planet. Arriving at the zoo at the beginning of the 20th cen…

(Seoul = Yonhap News) Reporter Kwak Min-seo = A famous tortoise at the San Diego Zoo in the United States, born in the 19th century, has passed away at the age of 141.

·Korea, Republic of
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Los Angeles (USA), Nov 25 (EFE).- Gramma, the Galapagos turtle nicknamed the “Zo zoo queen” of San Diego, died at 141 years old, reported the entity through its social networks. “For almost a century, Gramma, the Galapagos turtle, has been a silent and constant presence in the Zoo, witness to history, a beloved icon and an extraordinary ambassador for her species. With immense sadness, we share her death on November 20”, read in the announcement…

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Times of San Diego broke the news in San Diego County, United States on Friday, November 21, 2025.
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